Francois Clarac

post doctoral at Department of Organismal Biology, Evolution och utvecklingsbiologi

Visiting address:
Evolutionsbiologiskt Centrum EBC Norbyv. 18A
Postal address:
Norbyv. 18A
752 36 UPPSALA

Keywords: evolutionary biology paleobiology functional morphology herpetology

Also available at

My courses


  • PhD dissertation (2017):

The pseudosuchian dermal bone ornamentation: Morphology, Evolution, Functions.

  • Publication list:

1. F. Clarac, V. de Buffrénil, J. Cubo, and A. Quilhac. The ornamented osteoderm vascularization: Physiological implications in ectothermy and amphibious lifestyle in the crocodylomorphs? (2018) The Anatomical Record 301: 175-183.

2. F. Clarac, F. Goussard, V. de Buffrénil and V. Sansalone. Do the ornamented osteoderms influence the heat conduction through the skin? A finite element analysis in Crocodylomorpha. (2017) Journal of Thermal Biology 69: 39-53.

3. F. Clarac, V. de Buffrénil, C. Brochu and J. Cubo. The evolution of bone ornamentation in Pseudosuchia: morphological constraints versus ecological adaptations. (2017) Biological Journal of the Linnean Society 121: 395-408.

4. J. Cubo, M. Hui, F. Clarac and A. Quilhac. Static osteogenesis does not precede dynamic osteogenesis in periosteal ossification of Pleurodeles (Caudata, Amphibia) and Pogona (Squamata, Lepidosauria). (2017) Journal of Morphology 278(5): 621-628.

5. F. Clarac, T. Souter, J. Cubo, C. Brochu, V. de Buffrénil, R. Cornette. Does skull morphology constrain bone ornamentation? A morphometric analysis in the Crocodylia (2016) Journal of Anatomy 229: 292-301.

6. V. de Buffrénil, F. Clarac, A. Canoville, M. Laurin. Comparative data on the differentiation and growth of bone ornamentation in gnathostomes (Chordata: Vertebrata) (2016) Journal of Morphology 277(5): 634-670.

7. F. Clarac, T.Souter, R.Cornette, J.Cubo, V. de Buffrénil. A quantitative assessment of bone area increase due to ornamentation in the Crocodylia (2015) Journal of Morphology 276: 1183-1192.

8. V. de Buffrénil, F. Clarac, M. Fau, S. Martin, B. Martin, E. Pelle, and M. Laurin. Differentiation and growth of bone ornamentation in vertebrates: A comparative histological study among the Crocodylomorpha (2015) Journal of Morphology 276: 425-445.

9. N. Rabet, F. Clarac, P. Lluch, E. Gallerne, M. Korn. Review of the Eulimnadia (Branchiopoda: Spinicaudata) from North Africa and adjacent regions, with two new species from Mauritania (2015) Journal of the Crustacean Biology 35(3): 461-472.


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